Publications
Amélineau, F., Grémillet, D., Harding, A.M.A., Walkusz, W., Choquet, R., and Fort, J. 2019. Arctic climate change and pollution impact little auk foraging and fitness across a decade. Scientific Reports volume 9, Article number: 1014
Amélineau, F., Bonnet, D., Heitz, O., Mortreux, V., Harding, A.M.A., Karnovsky, N., Walkusz, W., Fort, J., and Grémillet, D. 2016. Microplastic pollution in the Greenland Sea: Background levels and selective contamination of planktivorous diving seabirds. Environmental Pollution 219:1131-1139. doi: 10.1016/j.envpol.2016.09.017.
Wojczulanis-Jakubus, K., Kilikowska.A., Harding, A.M.A., Jakubas, D., Karnovsky., N.J., Steen, H., Strøm, Welcker, J., Gavrilo, M., Lifjeld, J.T., and Johnson, A. (2014). Weak population genetic differentiation in the most numerous Arctic Seabird, the Little Auk. Polar Biology Volume 37 (5) pp 621–630
Parades, R., Orben, R.A., Suryan, R.M., Irons, D.B., Roby, D.D., Harding, A.M.A., Young, R.C., Beniot-Bird, K., Ladd, C., Renner, H., Heppel, S., Phillips, R.A., and Kitaysky, A. (2013) Foraging Responses of black-legged kittiwakes to prolonged food-shortages around colonies in the Bering Sea Shelf. PLoS ONE 9(3): e92520. https://doi.org/10.1371/journal.pone.0092520
Paredes, R., Harding, A.M.A., Irons, D.B., Roby, D.D., Suryan, R.M., Orben, R.A., Renner, H., Young, R., and Kitaysky, A., 2012. Proximity to multiple foraging habitats enhances seabirds’ resilience to local food shortages. Marine Ecology Progress Series.
Brown, Z.A., Welcker, J., Harding, A.M.A., Walkusz, W., and Karnovsky, N.J. (2012) Divergent diving behavior during short and long trips of a bimodal forager, the little auk Alle alle. Journal of Avian Biology 43: 215-226.
Gremillet, D., Welcker, J., Karnovsky, N.J., Walkusz, W., Hall, M.E., Fort, J., Brown, Z.W., Speakman, J.R. & Harding, A.M.A. (2012). Little auks buffer the impact of current Arctic climate change. Marine Ecology Progress Series 454: 197-206.
Mosbech, A., Johansen, K.L., Bech, N.I., Lyngs, P, Harding, A.M.A., Egevang, C., Phillips, R.A., and Fort, J. (2012) Inter-breeding movements of little auks Alle alle reveal a key post-breeding staging area in the Greenland Sea. Polar Biology 35: 305-311.
Karnovsky, N.J., Brown, Z.W., Welcker, J., Harding, A.M.A., Walkusz, W., Cavalcanti, A., Hardin, J., Kitaysky, A., Gabrielsen G., and Grémillet, D. (2011) Inter-colony comparison of diving behavior of an Arctic top predator: implications for warming in the Greenland Sea. Marine Ecology Progress Series: 10.3354/meps09351.
Harding, A.M.A., Welcker, J., Steen, H., Hamer, K.C., Kitaysky, A.S., Fort, J., Talbot, S.L., Cornick, L.A., Karnovsky, N.J., Gabrielsen, G.W., and Grémillet, G. (2011) Adverse foraging conditions may impact body mass and survival of a high Arctic seabird. Oecologia 167: 49-59.
Wojczulanis-Jakubas, K., Jakubas, D., Welcker, J., Harding, A.M.A., Karnosky, N.J., Kidawa, D., Steen, H., Stempniewicz, L. and Camphuysen, C.J. (2010) Body size variation of a high-Arctic seabird: the dovekie (Alle alle). Polar Biology 34: 847-854.
Fort, J., Cherel, Y., Harding, A.M.A., Egevang, C., Steen, H., Kuntz, G., Porter, W.P., and Grémillet, D. (2010) The feeding ecology of little auks raises questions about winter zooplankton stocks in North Atlantic surface waters. Biology Letters 6: 682-684.
Fort, J., Cherel, Y., Harding, A.M.A., Welcker, J., Jakubas, D., Steen, H., Karnovsky, N.J., and Grémillet, D. (2010) Geographic and seasonal variability in the isotopic niche of little auks. Marine Ecology Progress Series 414: 293-302.
Karnovsky, N., Harding, A.M.A., Walkusz, V., Kwaśniewski, S., Goszczko, I., Wiktor, J., Routti, H., Bailey, A., McFadden, L., Brown, Z., Beaugrand, G., and Grémillet, D. (2010) Foraging distributions of little auks (Alle alle) across the Greenland Sea: Implications of present and future climate change. Marine Ecology Progress Series 415: 283-293.
Shultz, M.T., Piatt, J.F., Harding, A.M.A., Kettle, A.B., and Van Pelt, T.I. (2009). Timing of breeding and reproductive performance in murres and kittiwakes reflect mismatched seasonal prey dynamics. Marine Ecology Progress Series 393: 247-258.
Harding, A.M.A., Kitaysky, A.S., Hamer, M.E., Hall, M.E., Welcker, J., Talbot, S.L., Karnovsky, N.J., Gabrielsen, G.W., and Grémillet, D. (2009) Impacts of experimentally increased foraging effort on the family: offspring sex matters. Animal Behaviour 78: 321-328.
Welcker, J., Harding, A.M.A., Kitaysky, A.S., Speakman, J.R., and Gabrielsen, G.W. (2009) Daily energy expenditure increases in response to low nutritional stress in an Arctic-breeding seabird with no effect of mortality. Functional Ecology 107: 1-97.
Welcker, J., Steen, H., Harding, A.M.A., and Gabrielsen, G.W. (2009) Sex-specific provisioning behavior in a monomorphic seabird with a bimodal foraging strategy. Ibis 151: 502-513.
Harding, A.M.A., Egevang, C., Walkusz, W., Merkel, F., Blanc, S., and Grémillet.D. (2009) Estimating prey capture rates of a planktivorous seabird, the little auk (Alle alle), using diet, diving behaviour, and energy consumption. Polar Biology 32: 785-796.
Welcker, J., Harding, A.M.A., Karnovsky, N.J., Steen, H., Strøm, H., and Gabrielsen, G.W. (2009) Flexibility in the bimodal foraging strategy of a high Arctic marine predator. Journal of Avian Biology 40 (4): 388-399.
Harding, A.M.A., Kitaysky, A.S., Hall, M.E., Welcker, J., Karnovsky, N.J., Talbot, S.L., Hamer, K.C., and Grémillet, D. (2009) Flexibility in the parental effort of an Arctic-breeding seabird. Functional Ecology 23: 348-358.
Harding, A.M.A, Hobson, K.H., Wojciech, W., Dmoch, K, Karnovsky, N, Van Pelt, T.I., and Lifjeld, J. (2008) Can stable isotope (δ13C and δ15N) measurements of little auk (Alle alle) adults and chicks be used to track changes in high-Arctic marine foodwebs? Polar Biology 31 (6): 725-733.
Harding, A.M.A., Piatt, J.F., and Schmutz, J.A. (2007) Seabird behaviour as an indicator of food supplies: Sensitivity across the breeding season. Marine Ecology Progress Series 352: 269-274.
Harding, A.M.A., Piatt, J.F., Schmutz, J.A., Kettle, A.B., Shultz, M., Van Pelt, T.I., Kettle, A.B., and Speckman, S.G. (2007) Prey density and the behavioural flexibility of a marine predator: the common murre (Uria aalge). Ecology 88 (8): 2024-2035.
Piatt, J.F., Harding, A.M.A., Shultz, M., Speckman, S.G., Van Pelt, T.I., Drew, G.S., and Kettle, A.B. (2007) Seabirds as indicators of marine food supplies: Cairns revisited. Marine Ecology Progress Series 352: 221-234.
Renner, H.M., Renner, M., Harding, A.M.A., Jones, I.J., Irons, D.B., Reynolds, J.H., and Byrd, G.V. (2006) Auklet population monitoring by colony mapping: results from St. George Islands, Alaska. Condor 108: 423-434.
Harding, A.M.A., Piatt, J.F., Byrd, G.V., Hatch, S.A., Konyukhov, N.B., and Golubova, E.U. (2005) Journal of Wildlife Management. Variability in colony attendance of crevice nesting Horned Puffins: implications for population monitoring. Journal of Wildlife Management 69 (3): 1279-1296.
Lifjeld, J.T., Harding, A.M.A., Mehlum, F., and Øigarden, T. (2005) No evidence of extra-pair paternity in the Little Auk (Alle alle). Journal of Avian Biology 36: 484-487
Harding, A.M.A., Van Pelt, T.I., Lifjeld, J.T., and Mehlum, F. (2004) Sex differences in Little Auk Alle alle parental care: transition from biparental to paternal-only care. Ibis 146: 642-651.
Harding, A.M.A., Piatt, J.F., and Hamer, K.C. (2003) Breeding ecology of horned puffins in Alaska: annual variation and effects of El Niño. Canadian Journal of Zoology 81: 1004-1013.
Harding, A.M.A., Van Pelt, T.I., Piatt, J.F., and Kitaysky, A.S. (2002) Reduction of provisioning effort in response to experimental manipulation of chick nutritional status in the horned puffin Fratercula corniculata. The Condor 104: 842-847.
Book Chapters and POPULAR READING
Piatt, J.F., and A.M.A. Harding 2007. Population Ecology of Seabirds in Cook Inlet. Pp. 335-352 in: Robert Spies (ed.), Long-term Ecological Change in the Northern Gulf Of Alaska. Elsevier, Amsterdam. In press (Dec. 13, 2006)
Suluk 2005 #5. Air Greenland in-flight magazine. “Greenland is the realm of the dovekie” Carsetn Evegang, Ann Harding, and Flemming Merkel.
Arctic Bulletin 4.05. 2005. Auk Clue to Climate Change. Ann Harding, David Grémillet, Nina Karnovsky and Geir Wing Gabrielsen.
Reports
Harding, A.M.A., Barger, C., Will, A. and Papish, R. (2009) Bogoslof 2009 Patch Dynamics Study. Unpublished USFWS report, AMNWR 09/02. Homer Alaska.
Piatt, J. F., Kuletz, K.J., Burger, A.E., Hatch, S.A., Friesen, V.L., Birt, T.P., Arimitsu, M.L., Drew, G.S., Harding, A.M.A. and Bixler, K.S. (2006) Status Review of the Marbled Murrelet (Brachyramphus marmoratus) in Alaska and British Columbia: U.S. Geological Survey. Open-File Report 2006-1387, 630pp.
Piatt, J., Sydeman, W., Harding, A., Byrd, V., Irons, D., Springer, A., and Kuletz, K. (2006) Seabirds as Indicators of Marine Ecosystems: An Integrated NPRB Science Plan for Alaska. Final Report NPRB Project 516.
Piatt, J.F., Byrd, G.V., Litzow, M.A., Shultz, M., Van Pelt, T., Kettle, A., and Harding, A.M.A. (2004) Protocols for long-term monitoring of seabird ecology in the Gulf of Alaska. Exxon Valdez Oil Spill Restoration Project Final Report (Restoration Project 00501), U.S. Geological Survey, Anchorage, Alaska.
Harding, A.M.A. (2004) Least Auklet Adult Survival and Colony Mapping. Alaska Maritime National Wildlife Refuge report 03/14: Rev. 1.
Harding, A.M.A., and Guldager, N. (2004) Least Auklet Nest Boxes, St. George, Pribilofs, 2003. Alaska Maritime National Wildlife Refuge report 03/14.
Harding, A.M.A., Piatt, J.F., Byrd, G.V., Hatch, S.A., Konyukhov, N.B., and Golubova, E.U. 2002. Attendance patterns and population monitoring of crevice-nesting Horned Puffins and Parakeet Auklets in the North Pacific. Final Rep. to USGS Species at Risk Prog. (Washington DC). USGS Alaska Science Center, Anchorage, AK. 50 pp.
Piatt, J. F. (ed.). (2002) Response of seabirds to fluctuations in forage fish density. Final Report to Exxon Valdez Oil Spill Trustee Council (Restoration Project 00163M) and Minerals Management Service (Alaska OCS Region). Alaska Science Center, U.S. Geological Survey, Anchorage, Alaska.
Zador, S., A. Harding, J.F. Piatt, L. Ochikubo and A. Neilsen. (1997) Monitoring Populations and Productivity of Seabirds at Colonies in Lower Cook Inlet, Alaska 1995. U.S. Geological Survey. Final report for Minerals Management Service (OCS Study MMS 95-0063). Anchorage, Alaska. 50 pp.
Harding, A. (1994) Common Murre on the menu. Pacific Seabirds, Vol. 21, No. 2.
Harding, A., Paredes, R., Suryan, R., Roby, R., Irons, D., Orgen, R., Renner, H., Young, R., Barger, C., Dorresteijn, I., and Kitaysky, A., (2013). Does location really matter? An inter-colony comparison of seabirds breeding at varying distances from productive oceanographic features in the Bering Sea. Deep Sea Research Pat II: Tropical Studies in Oceanography 94: 178-191
Harding, A., N. Fomina, O. Belonovich, A. Barsukova, T. Kushin, R. Papish, A. D. Lestenkof, K. Bush, H. Renner, K. Holser, and P. Wohl. (2021). A thousand miles of islands: Building relationships between young people of the Commander and Pribilof islands. Alaska Park Science 20(2): 52-57.
